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The absence of air sacs in humans may hold clues to speech evolution, but little is known about their functions in extant apes. The two series of grunts used in our analysis were the only ones that allowed us to confidently distinguish between the series of the male and female, which was necessary to reliably measure their pulse frequencies. Information about the distance of the subject from the microphone would facilitate synchrony analysis. This is also consistent with anecdotal reports that male orangutans inflate their sacs as part of a visual size exaggeration display Marler and Tenaza ; Tuttle Duration and pulse mean and range frequency pps: Thus our findings may reflect genus-wide vocal behavior.